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A private person (non FFL holder) can ship a rifle by any method...UPS, USPS, FedEx...to an FFL. A private person can ship a handgun via UPS or FedEx to an FFL. There is no requirement that a pistol be shipped by an FFL...only that it be received by an FFL. Just like there's no law that it be sent via air by UPS or FedEx. Both shippers require it, but not based on any legal requirement.A rifle can be taken directly to FEDEX etc to a FFL, but a pistol has to go FFL to FFL.
There are some exceptions to this. For example, if you're going hunting in Colorado you could ship your rifle to your buddy's house and take possession of it once there. But those are the general rules.